for his work on ribonucleaseandfor their contribution to the understanding of the connection between chemical structure and catalytic activity of the ribonuclease molecule
for their elucidation of the enzymatic mechanism underlying the synthesis of adenosine triphosphateandfor his discovery of an ion-transporting enzyme, Na+, K+ -ATPase
... Joanna of Castile and grandson of Ferdinand II of Aragon and Isabella I of Castile and of Maximilian I, Holy Roman Emperor.
Charles was born in Ghent and brought up in ...