for his work on ribonucleaseandfor their contribution to the understanding of the connection between chemical structure and catalytic activity of the ribonuclease molecule
for their elucidation of the enzymatic mechanism underlying the synthesis of adenosine triphosphateandfor his discovery of an ion-transporting enzyme, Na+, K+ -ATPase
... similar to those found in mitochondria), and then plastocyanin[?] before returning to chlorophyll. This transport chain produces a proton-motive force, pumping H+ ions ...