for his work on ribonucleaseandfor their contribution to the understanding of the connection between chemical structure and catalytic activity of the ribonuclease molecule
for their elucidation of the enzymatic mechanism underlying the synthesis of adenosine triphosphateandfor his discovery of an ion-transporting enzyme, Na+, K+ -ATPase
... and morality.
The novel intially appeared serially in the periodical Ruskii Vestnik ("Russian Messenger"), but Tolstoy clashed with the editor, Mikhail Katkov, over ...